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Huxley, Thomas Henry, 1825-1895

"American Addresses, with a Lecture on the Study of Biology"


What is commonly called the knee of a horse is its wrist. The "cannon
bone" answers to the middle bone of the five metacarpal bones, which
support the palm of the hand in ourselves. The "pastern," "coronary,"
and "coffin" bones of veterinarians answer to the joints of our middle
fingers, while the hoof is simply a greatly enlarged and thickened nail.
But if what lies below the horse's "knee" thus corresponds to the middle
finger in ourselves, what has become of the four other fingers or
digits? We find in the places of the second and fourth digits only two
slender splint-like bones, about two-thirds as long as the cannon bone,
which gradually taper to their lower ends and bear no finger joints, or,
as they are termed, phalanges. Sometimes, small bony or gristly nodules
are to be found at the bases of these two metacarpal splints, and it is
probable that these represent rudiments of the first and fifth toes.
Thus, the part of the horse's skeleton, which corresponds with that of
the human hand, contains one overgrown middle digit, and at least two
imperfect lateral digits; and these answer, respectively, to the third,
the second, and the fourth fingers in man.
Corresponding modifications are found in the hind limb.


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